Papers and others
〈Book chapter *as a corresponding author〉
2. Ushimaru A, Uchida K, Ikegami M and Suka T (2020)
Grasslands and shrublands in Japan. pp 785-799. in Goldstein MI & DellaSala L (eds.) . The Encyclopedia of the World's Biomes, Vol.3 section 6: Grasslands and shrublands - Sea of plants Elsevier, Oxford.
1. Ushimaru A*, Uchida K and Suka T (2018)
Grassland biodiversity in Japan: threats, management and conservation. pp 197-218. in Squires VR, Dengler J, Feng H & Hua L (eds.) 2018. Grasslands of the world: diversity, management and conservation CRC Press, Boca Raton, US. (All authors contributed equally to the paper)
〈Papers * as a corresponding author〉
107. Tomitaka M, Inoue T, Hirayama GH, Ushimaru A, Ishii HS, Sasaki T and Kenta T
(2025)
Aesthetic ecosystem services of diverse wildflowers: People's experience shapes preferences for floral traits. Nature-Based Solutions xx:xx-xx.
106. Watazu T, Hiraiwa MK, Inoue S, Mishima H, Ushimaru A and Hosaka T
(2025)
Effectiveness of interval photography cameras for a survey of pollinator communities: comparison with direct observation. Applications in Plant Sciences xx:xx-xx.
105. Uchiyama Y, Kyan A, Sato M, Ushimaru A, Minamoto T, Harada K, Takakura M, Kohsaka R, Kiyono M, Tsurumi T, Uchida A, Saga T and Yamamoto K
(2025)
Association between objective and subjective relatedness to nature and human well-being: Key factors for residents and possible measures for inequality in Japan's megacities. Landscape and Urban Planning 261:105377.
104. Ishii HS, Tsujimoto SG, Kadoya MB, Watanabe Y, Kohyama TI, Hiraiwa MK, Ushimaru A and Kudo G
(2025)
Linkage between community assemblage of floral traits and pollinator fauna: a comparison among six geographical regions. Plant Biology 27:538-548.
103. Watazu T, Hiraiwa MK, Ushimaru A and Hosaka T
(2025)
Pollinator is a housemate: Aeinetia indica, a parasite on Miscanthus sinensis, is pollinated by Ceratina bees nesting in its host. Plant Species Biology 40:266-273.
102. Hirayama GS, Inoue T, Kenta T, Ishii HS and Ushimaru A
(2025)
Long-term management is required for the recovery of pollination networks and function in restored grasslands. Journal of Applied Ecology 62:814-823.
101. Nammoku Y, Nikkeshi A, Terai Y, Ushimaru A and Kinoshita M
(2025)
Morphological and DNA qnalysis of pollen grains on the butterfly individuals reveal their flower visitation history. The Science of Nature-Naturwissenschaften 112:13.
100. Asada AA*, Ohwaki A, Yaida YA, Kawakami F, Masuda M and Ushimaru A*
(2025)
Prescribed burning effectively maintains threatened species in semi-natural grasslands on lava flows Plants, People, Planet xx:xx. (AAA and AU contributed equally to the paper)
99. Hirayama GS, Tomita M and Ushimaru A
(2025)
Impacts of agricultural intensification and abandonment on plant and pollinator diversity, their mutualistic networks and pollination services Agriculture, Ecosystems & Environment 381:109437. (GSH and AU contributed equally to the paper)
98. Shimomai H, Taichi N, Katsuhara KR, Kato S, Ushimaru A, and Ohmido N
(2024)
Allopolyploidy enhances survival advantages for urban environments in the native plant genus Commelina. Annals of Botany 134:1055-1066.
97. Uchiyama Y, Kyan A, Sato M, Ushimaru A, Minamoto T, Kiyono M, Harada K and Takakura M
(2024)
Local environment perceived in daily life and urban green and blue space visits: uncovering key factors for different age groups to access ecosystem services Journal of Environmental Management 370:122676.
96. Hirayama GH and Ushimaru A
(2024)
Habitat preference influences response to changing argicultural landscapes in two long-horned bees. Journal of Apicultural Research 63:1067-1077.
95. Katsuhara KR, Ushimaru A and Miyazaki Y
(2024)
Does coexisting congener of a mixed mating species affect the genetic structure and selfing rate via reproductive interference? Oecologia 206:37-45.
94. Yaida YA*, Nagai T, Oguro K, Katsuhara KR, Uchida K, Kenta T and Ushimaru A*
(2024)
Negative legacy effects of past forest use on plant diversity in semi-natural grasslands on ski slopes Journal of Environmental Management 368:122120. (YAY, TK & AU contributed equally to the paper)
93. Nakahama N, Konagaya T, Ueda S, Hirai N, Yago M, Yaida YA, Ushimaru A, and Isagi Y
(2024)
Road to extinction: archival samples unveiled the process of inbreeding depression during artificial breeding in an almost extinct butterfly species. Biological Conservation 296:110686.
92. Yaida YA*, Inoue T, Kenta T and Ushimaru A*
(2024)
Seed dispersal limitation causes negative legacy effect on restoration of grassland plant diversity on ski slopes. Ecology and Evolution 14:e11654.
91. Taichi N*, Nakahama N, Ohmido N and Ushimaru A*
(2024)
Habitat diversification associated with urban development has a little effect on genetic structure in the annual native plant Commelina communis in an East Asian megacity. Ecology and Evolution 14:e10975.
90. Hiraiwa MK and Ushimaru A
(2024)
Loss of functional diversity rather than species diversity of pollinators decreases community-wide trait matching and pollination function. Functional Ecology 38:1296-1308.
89. Watazu T, Hiraiwa MK, Inoue S, Mishima H, Ushimaru A and Hosaka T
(2024)
Dominance of non-wetland-dependent pollinators in a plant community in a small natural wetland in Shimane, Japan. Journal of Plant Research 137:191-201.
88. Hidaka S, Jo TS, Yamamoto S, Katsuhara KR, Tomita S, Miya M, Ikegami M, Ushimaru A and Minamoto T
(2024)
Sensitive and effective surveillance of Japanese giant salamander (Andria japonicus) distribution in western Japan using multi-copy nuclear DNA marker. Limnology 137:191-201.
87. Taichi N and Ushimaru A
(2024)
Trait variation along an urban-rural gradient in Asian dayflower: the contribution of phenotypic plasticity and genetic divergence. Plant Biology 26:74-81.
86. Nakahama N, Kurata S and Ushimaru A (2023)
Contribution of genetic analyses to semi-natural grassland biodiversity conservation in Japan. Plant Species Biology 38:158-170.
85. Sato M, Minamoto T and Ushimaru A
(2023)
An interdisciplinary approach to environmental conservation policy: a case of Satoyama re-development in the peri-urban area. International Journal of Economic Policy Studies 17:403-419.
84. Aoshima I, Nakao R, Minamoto T, Ushimaru A, Sato M
(2023)
Heterogenous preference for biodiversity in Japanese urban blue spaces based on people's nature experiences: analysis using eDNA and satisfaction data. City and Environment Interactions 18:100101.
83. Ushimaru A*, Seo N, Sakagami K and Funamoto
(2023)
Sexual dimorphism in dioecious species with complex, specialist pollinated flowers. American Journal of Botany 110:e16148. (AU and NS contributed equally to the paper)
82. Nishizawa R, Nakao R, Ushimaru A and Minamoto T
(2023)
Development of evironmental DNA detction assays for snakes in paddy fields in Japan. Landscape and Ecological Engineering 19:3-10.
81. Jo T, Sato M, Minamoto T and Ushimaru A
(2022)
Valuing the cultural services from urban blue space ecosystems in Japanese megacities during the COVID-19 pandemic. People and Nature 4:1176-1189.(All authors contributed equally to the paper)
80. Nakata T*, Ishii R, Yaida YA and Ushimaru A*
(2022)
Horizontal orientation facilitates pollen transfer and rain damage avoidance in actinomorphic flowers of Platycodon grandiflorous. Plant Biology 24:798-805.(TN & AU contributed equally to the paper)
79. Murakami K, Katsuhara KR and Ushimaru A
(2022)
Intersexual flower differences in an andromonoecious species: small pollen‐rich staminate flowers under resource limitation. Plant Biology 24:259-265.
78. Katsuhara KR, Tachiki Y, Iritani R and Ushimaru A (2021)
The eco-evolutionary dynamics of prior selfing rates promote coexistence without niche partitioning under conditions of reproductive interference. Journal of Ecology 109:3916-3928.
77. Biurrun I et al. (2021)
Benchmarking plant diversity of Palaearctic grasslands and other open habitats. Journal of Vegetation Science 32:e13050. (223 coauthors and AU is 199th)
76. Ushimaru A*, Ishii R and Katsuhara KR
(2021)
Covering and shading by neighbouring plants diminish pollinator visits to and reproductive success of a forest edge-specialist dwarf species. Plant Biology 23:711-718. (AU & RI contributed equally to the paper)
75. Nikkeshi A, Hiraiwa MK, Ushimaru A, Hoshizaki K, Makita A and Inoue M (2021)
Evaluation of sampling effort sufficient to assess pollen species richness on pollinators using rarefaction. Applications in Plant Sciences 9:e11411.
74. Inoue T, Yaida Y, Uehara Y, Katsuhara KR, Kawai J, Takashima K Ushimaru A and Kenta T (2021)
The effects of temporal continuities of grasslands on the diversity and species composition of plants. Ecological Research 36:24-31.
73. Jo T, Tomita S, Kohmatsu Y, Osathanunkul M, Ushimaru A and Minamoto T
(2020)
Seasonal monitoring of Hida salamander Hynobius kimurae using environmental DNA with a genus-specific primer set. Endangered Species Research 43:341-352.
72. Iwachido Y, Uchida K, Ushimaru A Yokota S and Sataki T
(2020)
Nature-oriented park use of satoyama ecosystems can enhance biodiversity conservation in urban landscapes. Landscape and Ecological Engineering 16:163-172.
71. Jo T, Fukuoka A, Uchida K, Ushimaru A and Minamoto T
(2019)
Multiplex real-time PCR enables the simultaneous detection of environmental DNA from freshwater fishes: a case study of three exotic and three threatened native fishes in Japan. Biological Invasions 22:455-471.
70. Matsuhisa S and Ushimaru A
(2019)
Does sexual dimorphism exist in flowering phenology traits in anemophilous dioecious species?: a test with Rumex acetosa. American Journal of Botany xx:xx-xx.
69. Valko O, Labadessa R, Palpurina S, Burrascano S, Ushimaru A and Venn S
(2019)
Conservation and diversity of Palearctic grasslands - Editorial to the 5th EDGG special issue in Hacquetia. Hacquetia 18:143-146.
68. Katsuhara KR, Nakahama N, Komura T, Kato M, Miyazaki Y, Isagi Y, Ito M and Ushimaru A (2019)
Development of microsatellite markers for the annual andromonoecious herb Commelina communis f. ciliata (Commelinaceae). Genes & Genetic Systems 94: 133-138.
67. Yaida Y, Nagai T, Oguro K, Katsuhara KR, Uchida K, Kenta T and Ushimaru A (2019)
Ski runs as an alternative habitat for threatened grassland plant species in Japan. Palaearctic Grasslands 42: 16-22.
66. Katsuhara KR and Ushimaru A (2019)
Prior selfing can mitigate the negative effects of mutual reproductive interference between coexisting congeners. Functional Ecology 33: 1504-1513.
65. Ollerton J et al. (2019)
The diversity and evolution of pollination systems in large plant clades: Apocynaceae as a case study. Annals of Botany 123:311-325.(75 coauthors and AU is 69th)
64. Aoshima I, Uchida K, Ushimaru A and Sato M (2018)
The influence of subjective perceptions on the valuation of green spaces in Japanese urban areas. Urban Forestry & Urban Greening 34:166-174.
63. Dengler J et al. (2018)
GrassPlot-a database for multi-scale plant diversity in Palaearctic grasslands. Phytocoenologia 48:331-347. (183 coauthors and AU is 163rd)
62. Uchida K, Fujimoto H and Ushimaru A (2018)
Urbanization promotes the loss of seasonal dynamics in the semi-natural grasslands of the East Asian megacity. Basic and Applied Ecology 29: 1-11.
61. Nakahama N, Uchida K, Ushimaru A and Isagi Y (2018)
Historical changes in grassland area determined the demography of semi-natural grassland butterflies in Japan. Heredity 121:151-168.
60. Sato M, Ushimaru A and Minamoto T (2017)
The effect of different personal histories on valuation of forest ecosystem services in urban areas: a case study of Mt Rokko,Kobe, Japan.Urban Forestry & Urban Greening 28: 110-117.
59. Matsuhisa S, Yanagisawa N, Ushimaru A (2017)
Sexual dimorphism in vegetative traits in dioecious Ilex crenata
. GIFUKEN SHOKUBUTSUKENKYUKAISHI 33:1-7. (in Japanese)
58. Aoshima I, Uchida K, Ushimaru A, Sato M (2017)
Economic Valuation of the urban green spaces using life satisfaction data. Environmental Science 30:238-249. (in Japanese with English abstract)
57. Katsuhara KR, Kitamura S and Ushimaru A (2017)
Functional significance of petals as landing sites in fungus-gnat-pollinated flowers of Mitella pauciflora (Saxifragaceae). Functional Ecology 31: 1193-1200. (All authors contributed equally to the paper)
55. Hiraiwa MK and Ushimaru A (2017)
Low functional diversity promotes niche changes in natural island pollinator communities. Proceedings of the Royal Society B 284: 20162218.
55. Fukuoka A, Takahara T, Matsumoto M, Biology Club of Hyogo Prefectural Agricultural High School, Ushimaru A and Minamoto T (2016)
Establishment of detection system for native rare species, Hemigrammocypris rasborella, using environmental DNA. Japanese Journal of Ecology 66:613-620. (in Japanese with English abstract)
54. Nikkeshi A, Hiraiwa MK, Ushimaru A, Hoshizaki K, Makita A and Inoue M (2016)
Established method of deposited pollen grains on the surface of pollinated insect body. Japanese Journal of Palynology 62: 3-7. (in Japanese with English abstract)
53. Nagata YK* and Ushimaru A* (2016)
Traditional burning and mowing practices support high grassland plant diversity by providing intermediate levels of vegetation height and soil pH. Applied Vegetation Science 19: 567-577. (Both authors contributed equally to the paper)
52. Uchida K, Hiraiwa MK and Ushimaru A (2016)
Plant and herbivorous insect diversity loss are greater than null model expectations due to land-use changes in agro-ecosystems. Biological Conservation 201:270-276.
51. Uchida K, Takahashi S, Shinohara Tand Ushimaru A (2016)
Threatened herbivorous insects maintained by long-term traditional management practices in semi-natural grasslands. Agriculture, Ecosystems & Environment 221:156-162.
50. Nakahama N, Uchida K, Ushimaru A and Isagi Y (2016)
Timing of mowing influences genetic diversity and reproductive success in endangered semi-natural grassland plants. Agriculture, Ecosystems & Environment 221:20-27.
49. Nikkeshi A, Kurimoto D and Ushimaru A (2015)
Low flower-size variation in bilaterally symmetrical flowers: a support for the pollination precision hypothesis. American Journal of Botany 102:2032-2040 (All authors contributed equally to the paper)
48. Uchida K, Shinohara T, Takahashi S, Nakahama N, Takami Y and Ushimaru A
(2016)
Rediscovery of Celes akitanus (Orthoptera, Acrididae) from semi-natural grasslands in Japan. Entomological Science 19:89-96.
46. Kondoh M, Mougi A, Ushimaru A, and Nakata K
(2015)
Adaptive movement and food-chain dynamics. Theoretical Ecology 9:15-25.
46. Matsuhisa S and Ushimaru A
(2015)
Sexual dimorphism in floral longevity and flowering synchrony in relation to pollination and mating success in three dioecious Ilex species. American Journal of Botany 102:1187-1197.
45. Uchida K and Ushimaru A
(2015)
Land abandonment and intensification diminish spatial and temporal β-diversity of grassland plants and herbivorous insects within paddy terraces. Journal of Applied Ecology 52:1033-1043.
44. Ohara RG and Ushimaru A*(2015)
Plant beta-diversity is enhanced around grassland-forest edges within a traditional agricultural landscape. Applied Vegetation Science 18: 493-502.
43. Fukumoto S, Ushimaru A and Minamoto T
(2015)
A basin-scale application of enivironmental DNA assessment for rare endemic species and closely related exotic species in rivers: a case study of giant salamanders in Japan. Journal of Applied Ecology 52:358-365.
42. Uchida K and Ushimaru A
(2014)
Biodiversity declines due to abandonment and intensification of agricultural lands: patterns and mechanisms. Ecological Monographs 84:637-658.
41. Osawa T, Mitsuhashi H and Ushimaru A
(2014)
Plant species coexistence relationships may shift according to life histroy traits and seasons. Plant Ecology 215:597-612.
40. Ushimaru A*, Kobayashi A and Dohzono I
(2014)
Does urbanization promote floral diversification? : Implications from changes in herkogamy with pollinator availability in an urban-rural area. The American Naturalist 184:258-267. (AU & AK contributed equally to the paper)
39. Nakahama N, Ushimaru A and Isagi Y
(2013)
Reproductive traits and flower visitor fauna of Vincetoxicum pycnostelma Kitag. in the Nishitani area of Takarazuka City, Hyogo Prefecture. Bulletin of Kansai Organization for Nature Conservation 35 : 115-123. (in Japanese with English Abstract)
38. Uematsu Y and Ushimaru A*
(2013)
Topography- and management-mediated resource gradients maintain rare and common plant diversity around paddy terraces. Ecological Applications 23:1357-1366.
37. Nakata K and Ushimaru A
(2013)
The effect of predation risk on spider's decisions on web-site relocation. Behaviour 150:103-114.
36. Fukada C, Kaneko S, Yokogawa M, Kobayashi T, Ushimaru A and Isagi Y
(2012)
Development of ten microsatellite markers for Arisaema minus (Araceae), a vulnerable Japanese herb species. Conservation Genetics Resources 4:495-497.
35. Iwata T, Nagasaki O, Ishii HS and Ushimaru A*
(2012)
Inflorescence architecture affects pollinator behaviour and mating success in Spiranthes sinensis (Orchidaceae). New Phytologist 193:196-203.
34. Tsuji M, Ushimaru A*, Osawa T and Mitsuhashi H
(2011)
Paddy-associated frog declines via urbanization: a test of the dispersal-dependent-decline hypothesis. Landscape and Urban Planning 103:318-325.
33. Osawa T, Mitsuhashi H, Uematsu Y and Ushimaru A
(2011)
Bagging GLM: improved generalized linear model for the analysis of zero-inflated data. Ecological Informatics 6:270-275.
32. Osawa T, Mitsuhashi H, Niwa H and Ushimaru A
(2011)
The role of river confluences and meanderings in preserving local hot spots for threatened plant species in riparian ecosystems. Aquatic Conservation: Marine and Freshwater Ecosystems 21:358-363.
31. Ushimaru A* and Genkai-Kato M
(2011)
A theoretical framework for resource translocation during sexual reproduction in modular organisms. Evolutionary Ecology 25 :885-898.
30. Uematsu Y, Takeda Y and Ushimaru A
(2010)
The flora of Satokusachi in the Nishitani area of Takarazuka City, Hyogo Prefecture. Bulletin of Kansai Organization for Nature Conservation 32 :85 - 98. (in Japanese with English Abstract)
29. Osawa T, Mitsuhashi H, Niwa H and Ushimaru A
(2010)
Differences in distribution patterns around river confluences among hydrophilic vegetation groups. Ecological Research 25 :1161-1169.
28. Osawa T, Mitsuhashi H, Niwa H and Ushimaru A
(2010)
Enhanced diversity at network nodes: river confluences increase vegetation-patch diversity. The Open Ecology Journal 3 :48-58.
27. Osawa T, Mitsuhashi H and Ushimaru A
(2010)
River confluences enhance riparian plant species diversity. Plant Ecology 209 :95-108.
26. Tsujino R, Fujita N, Katayama M, Kawase D, Matsui K, Seo A,
Shimamura T, Takemon Y, Tsujimura N, Yumoto T and Ushimaru A (2010)
Restoration of floating mat bog vegetation after eutrophication damages by improving water quality in a small pond. Limnology 11 : 289-297.
25. Uematsu Y, Koga T, Mitsuhashi H and Ushimaru A
(2010)
Abandonment and intensified use of agricultural land decrease habitats of rare herbs in semi-natural grasslands. Agriculture, Ecosystems & Environment 135 :304-309.
24. Ushimaru A*, Dohzono I, Takami Y and Hyodo F (2009)
Flower orientation enhances pollen transfer in bilaterally symmetrical flowers. Oecologia 160 :667-674.
23. Yonekura R,
Yamanaka H, Ushimaru A, Matsui K, Uchii K and Maruyama A (2009)
Allochthonous prey subsidies provide an asymmetric growth benefit to invasive bluegills over native
cyprinids under the competitive conditions in a pond. Biological Invasions 11 : 1347-1355.
22. Ushimaru A*, Ishida C, Sakai S,
Shibata M, Tanaka H,
Niiyama K and Nakashizuka T (2008) The effects of human management on spatial
distribution of two bumble bee species in a traditional agro-forestry Satoyama landscape. Journal of Apicultural Research 47(4): 296-303.
21. Tsujino R,
Matsui K, Ushimaru A, Seo A, Kawase D, Uchihashi H, Kenji S,
Takahashi J, Yumoto T and Takemon Y (2007)
Invasion of the Mizorogaike Wetland by sika deer, and their effects on vegetation. Japanese Journal of Conservation Ecology 12: 20-27. (in Japanese with English Abstract)
20. Imamura
A and Ushimaru A (2007)
Flower orientation on slopes in myco-heterotrophic species, Monotropastrum globosum. Plant Species Biology 22: 161-166.
19. Ushimaru A*, Kikuchi S, Yonekura R,
Maruyama A, Yanagisawa N, Kagami M, Nakagawa M,
Mahoro S, Kohmatsu Y, Hatada A, Kitamura S and Nakata K (2007）The influence of floral
symmetry and pollination systems on flower size variation. Nordic Journal of Botany 24: 593-598.
18. Ushimaru A*, Watanabe T
and Nakata K. (2007) Colored floral organs
influence pollinator behavior and pollen transfer in Commelina communis. American Journal of Botany 94: 249-258.
17.
Ushimaru A*,
16.
Ushimaru A*
15. Nakata K and Ushimaru A (2004) Difference
in web construction behavior at newly occupied web sites between two Cyclosa species. Ethology.110:397-411.
14. Nakashizuka T et al. (2003) Monitoring dynamics of beech forests with different structure in Shirakami Mountains.
Tohoku Journal
of Forest Science 8:67-74. (in Japanese with English Abstract)
13. Nakata K and Ushimaru A and Watanabe T (2003) Using past experience in web relocation enhances the
foraging efficiency of the spider Cyclosa
argenteoalba. Journal
of Insect Behaviour 16:371-380.
12. Ushimaru A*, Itagaki T and Ishii
HS (2003) Floral correlations in an
andromonoecious Commelina communis. Plant Species Biology 18: 103-100
11. Ushimaru
A*, Itagaki T and Ishii HS (2003) Variation in floral organ size depends on function: a test
with Commelina communis, an
andromonoecious species. Evolutionary Ecology Research 5:615-622.
10. Ushimaru A*, Fukui A and Imamura A (2003) Effect of floral organ size on female reproductive
success in Erythronium japonicum
(Liliaceae). Journal of Plant Biology46: 245-249.
9. Watanabe T, Tanigaki T, Nishi H, Ushimaru
A and Takeuchi T (2002) Geographic color variation in two Geotrupes dung beetles: a further study.
Entomological
Science 5:291-295.
8. Ushimaru A* and Nakata K
(2002) The evolution of flower
allometry in selfing species. Evolutionary
Ecology Research 4:1217-1227.
7. Ushimaru A*
and Imamura A (2002) Large
flower size variation in the myco-heterotrophic plant, Monotropastrum globosum: effects of floral display on female
reproductive success. Plant Species Biology 17:
147-153.
6. Watanabe T, Tanigaki T, Nishi H, Ushimaru
A and Takeuchi T (2002) A
quantitative analysis of geographic color variation in two Geotrupes dung beetles.
Zoological Science 19:351-358.
5. Ushimaru A* and Nakata K
(2001) Evolution of flower
allometry and its significance on pollination success in deceptive orchid, Pogonia japonica. Internatinal Journal of Plant
Sciences 162:1307-1311.
4. Matsui K, Ushimaru A and Fujita
N (2001) Pollinator limitation in a deceptive
orchid, Pogonia japonica, on a
floating peat mat. Plant
Species Biology. 16:231-235.
3. Ushimaru A* and Matsui
K (2001) Sex change in tree species: long-term monitoring of sex
expression in Acer rufinerve. Nordic
Journal of Botany. 21:397-399.
2. Nakata K and Ushimaru A
(1999) Feeding experience affects
web relocation and investment in web threads in an orb-web spider, Cyclosa argeteoalba. Animal Behavior 57:1251-1255.
1. Ushimaru A* and Kikuzawa K (1999) Variation of breeding system, floral rewards, reproductive
success in clonal Calystegia
species. American
Journal of Botany 83:
436-446.
〈Others〉
5. Ushimaru A（2010）Floral evolution in zygomorphic flowers. Phenology Kenkyu 45:17-21. (in Japanese)
4. Ushimaru A（2009）Where do plants orient their flowers? : ecology of flower orientation. Seibutsukagaku 60:159-166. (in Japanese)
3. Ushimaru A（2009）Why do plants have so many flowers? Comments to the Miyadi Award paper by Ishii HS.
Japanese Journal of Ecology 58:170-172. (in Japanese)
2. Ushimaru A（2006）Lessons from floral guides. In: (edited by Kai S and Morikawa H) Plantmimetics, NTS Co.,
Tokyo, Pp 577-583. (in Japanese)
1. Ushimaru A（2000）Floral sex-Evolution of self-compatibility and autonomous
self-pollination in hermaphroditic flowering plants. In: (edited by SSSB) New Perspectives in Floral Ecology, Bun-ichi Sogo Shuppan Co.,
Tokyo, Pp 75-95 (in Japanese)
〈Book editing〉
1. Yuma M and Ushimaru A eds.（2007）Shinka no Katachi by Hokkaido University CoSTEP Science Writers.
Gijutsu-Hyohron Co., Ltd
〈Scientific reports〉
3. Ushimaru A, Ishida C, Sakai S, Shibata M, Tanaka H, Nagaike T, Niiyama K
and Nakashizuka T. (2008) Spatial distribution of two bumble bee species in an agro-forestry landscape. In Ichikawa M, Yamashita S and Nakashizuka T (eds)
“Sustainability and biodiversity
assessment on forest utilization options. ” Nakanishi Printing Company, Kyoto. Pp 221-226.
2. Kikuzawa K, Kimura K, Ushimaru A and Nomura N. (2002) Latitudinal and altitudinal gradients in species richness of
forest trees in relation to leaf, flower and fruit phenology. Reports of the MEXT Creative Basic
Research 09NP1501 “An
Integrated Study on Biodiverstiy Conservation under Global Change and
Bioinventory Management System” DIVER.
1. Endo A, Matsui K, Ushimaru Aand Fujita N (1996)
Characteristics of flower visitor community on a floating peat mat and secondary forests in Mizorogaike Pond.
Reports on animal community related to the road construction at Kamigamo, Kyoto City, Kyoto City Government, Kyoto, Japan. (in Japanese)
〈Theses〉
3. Ushimaru
A. 1998. Evolution of herkogamy and self-compatibility system in
flowering plants.
Kyoto University, Ph.D. thesis (Science) supervised by Kihachiro Kikuzawa
2. Ushimaru
A. 1995. Effects of self-incompatibility on fruit and seed set in
clonal Calystegia species. Kyoto University, M.S. thesis (Science) supervised by Hideo Tabata
1. Ushimaru A.
1992. Effect of storage
conditiions of seeds and water temperature during imbibition on flowering
responses in Pharbitis nil, strain Violet. Kyoto University, B.S. thesis (Agriculture) supervised by Masateru Shinozaki